Animal Corner 2003-2013. <>. Accessed 3 October 2013.

Arnaudin, M. E. 2012. Benefits of dung beetles (Colepltera: Scarabaeidae) on nutrient cycling and forage growth in alpaca pastures. Virginia Polytechnic Institute and State University.

Bertone, M.A. 2004. Dung beetles (Coleoptera: Scarabaeidae and Geotrupidae) Of North Carolina cattle pastures and their implications for pasture improvement. North Carolina State University.

Encyclopedia of Life 2013. <>. Accessed 3 December 2013.

Gittings, T. and P.S. Giller. 1997. Life history traits and resource utilisation in an assemblage
of north temperate Aphodius dung beetles (Coleoptera:Scarabaeidae). Ecography 20:55-66.

Holter, P. 1982. Resource utilization and local coexistence in a guild of Scarabaeid dung
beetles (Aphodius spp.). Oikos 39:213-217.

Holter, P. 1979. Effect of dung-beetles (Aphodius spp.) and earthworms on the disappearance of
cattle dung. Oikos 32:393-402.

Menendez, R, and D. Gutierrez. 1999. Heterotrophic succession within dung-inhabiting beetle
communities in northern Spain. Acta Oecologica 20:527-353.

Miller, A. 1961. The mouth parts and digestive tract of adult dung beetles (Coleptera:Scrarbaeidae) with reference to the ingestion of helminth eggs. The Journal of Parasitology 47: 735-744.

Nichols, E, S. Spector, J. Louzada, T. Larsen, S. Amezquita, and M.E. Favila. 2008. Ecological
functions and ecosystem services provided by Scarabaeinae dung beetles. Biological Conservation 141:1461-1474.

NSW Department of Primary Industries 2004. <>. Accessed 6 October 2013.

Pennington, J.A, K. VanDevender and J.A. Jennings. Nutrient and fertilizer value of dairy manure.University of Arkansas. Division of Agriculture. Cooperative Extension Service. Agriculture and Natural Resources: FSA4017.

Scarab for Kids 2009. <>. Accessed 3 December 2013.

Sladecek, F.X.J, J. Hrcek, P. Kilmes, and M. Konvicka. 2013. Interplay of succession and
seasonality reflects resource utilization in an ephemeral habitat. Acta Oecologica 46:17-24.

Stevenson, A. and C.A. Lindberg. 2010. New oxford American dictionary. Oxford University Press, New York, New York, USA.

Yoshida, N. and H. Katakura. 1986. Life cycles of Aphodius dung beetles (Scarabaeidae,
Coleoptera) in Sapporo, Northern Japan.  Graduate School of Environmental Science 8:209-229.

Young, O. P. 1998. Observations of rove beetles (Coleoptera: Straphylinidae) predation of dung beetles (Scarabaeidae) in Panama. The Coleopterists Bulletin 52: 217-222.