Chauliognathus pennsylvanicus is a dioecious species, which includes a definite separation between female and male sexes.  Soldier beetles reproduce sexually and reproduce 1 or 2 generations a year. (Swain 1948)  Most larvae feed on soft insects, like caterpillars, grasshopper eggs and maggots.  In the later stages of maturation, larvae spend winter pupating in soil cells underground. The larval environments usually consist of loose bark and damp ground or damp crevices. (White 1983) The larvae is often covered with a "fur" that looks similar to velvet, but has a dense texture. (White 1983)  The soldier beetles are most active during late summer and fall.                                 

Chauliognathus pennsylvanicus