Geography
According to The Distribution of the Native Land Molluscs of the Eastern United States  by the late Leslie Hubricht, the Upland Pillsnail populates the blackened countries on the map to the left (Hubricht 1985).  They can be found on the east side of the Great Smoky Mountains in the Big Creek Drainage, and in southeastern Ontario, Canada, as well (Grimm 2000; Dourson 2006).

Distribution of Upland Pillsnail. Hubricht, L. 1985. The Distribution of the Native Land Molluscs of the Eastern United States. —Fieldiana Zoology 24: 41. 

 Habitat
The habitat of Euchemotrema fraternum is terrestrial but also must be moist.  They are found under rocks and forest debris such as hardwood (non-coniferous tree) logs (Hubricht 1985). They have been found with other animals in old growth of forest with material from trees such as the American  Beech, American Hophornbeam, Spruce, Yellow Birch, Eastern Hemlock, and Sugar Maple (Hotopp & Smith 1995).  Because they are found in wooded areas, their populations are sensitive to urban sprawl as their amount of habitat is decreased.  Acid rain also takes a toll on these snails, and thus on the their predators and food sources.  Other things that affect how comfortable Upland Pillsnails are in their habitat are invasive species and forest fires.  Forest fires decrease the bottom layer of the forest where Upland Pillsnails live and obtain their food (Dourson 2006).  Upland Pillsnails have also been found to be arboreal (inhabitants of trees) in a study by Karen D. McCoy and Thomas D. Nudds called "Interspecific Variation in Climbing by Gastropods: Implication for Transmission of Pareolaphostrongylus tenius."  In this study, the Upland Pillsnail was found to be mostly arboreal and thus less likely to be host to that specific parasite, since they spend less time on the ground spreading the parasite to other animals (McCoy & Nudds 1997). 

To learn more about the effects of other species on the Upland Pillsnail, visit the Interactions page.