Not much is known about the diet of Daeadalochila uvulifera, but can be speculated that they are herbivorous.  Many of the Polygyrids are also mychophagus or fruiting fungi feeders (Perez, 2008).

Daedalochila uvulifera has a radula like all other gastropods with a few exceptions.  The radula is a rasping tonguelike organ (Hickman et al, 2009).  It is a ribbonlike tissue with chitinous teeth that point backward (Hickman et al, 2009).  The radula has a couple purposes, one is to scrape or rake up small bit of food, usually microscopic organisms or feed directly on plant matter (Hickman et al, 2009).  The food is then sent down the esophagus, where it then enters the stomach.  Daedalochila uvulifera has a complete digestive tract.  The digestive gland releases enzymes that breaks down the food.  After the nutrients have been absorbed, the waste is then excreted by the anus, the snail then sends to the nephridiopore with the help of wind or a current to prevent fouling, which can kill the snail (Heckman et al, 2009). 

Daedalochila uvulifera, has an open circulatory system, meaning they don't have capillaries (Heckman et al, 2009).  The heart is enclosed in a chamber, pumping hemolymph to the tissues of the body to supply oxygen (Heckman et al, 2009).  The hemolymph is contained in the sinuses surrounding the organs (Heckman et al, 2009).  An open circulatory system is less efficient in supplying tissue with the needed oxygen, so it is more common in slow moving animals (Heckman et al, 2009).